Irradiance-dependent opening and closure of stomata is comparably slow. Therefore, stomatal conductance (gs) often limits photosynthesis in fluctuating irradiance, particularly after large stepwise increases in irradiance in leaves adapted to low irradiance or darkness. This limitation to photosynthesis may reduce crop productivity in natural environments and greenhouses, where irradiance incident on a leaf can fluctuate rapidly. To test whether this limitation applies to typical greenhouse crops such as tomato (Solanum lycopersicum L.), photosynthetic gas exchange of flacca, a mutant with very high gs (2-4 times wildtype level) was compared with its wildtype (WT), cv. Rheinlands Ruhm. The steady-state response to leaf internal CO2 concentration (Ci) was similar between genotypes, indicating similar properties of the leaf photosynthetic apparatus. Surprisingly, when exposing dark-adapted leaves in ambient CO2 concentration (400 ppm) to a stepwise increase
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